(A) Centrophryne spinulosa , 136 mm SL (B) Cryptopsaras couesii , 34.5 mm SL (C) Himantolophus appelii , 124 mm SL (D) Diceratias trilobus , 86 mm SL (E) Bufoceratias wedli , 96 mm SL (F) Bufoceratias shaoi , 101 mm SL (G) Melanocetus eustalus , 93 mm SL (H) Lasiognathus amphirhamphus , 157 mm SL (I) Thaumatichthys binghami , 83 mm SL (J) Chaenophryne quasiramifera , 157 mm SL.
Udičarkе (Lophiiformes) su red riba iz infraklase pravih košljoriba (Teleostei). To je košljoriba imenovana po svom karakterističnom vidu predacije , u kome, u kome, mesnata izraslina iz glave ribe (esca ili illicium ) deluje kao mamac, koji privlači druge ribe.
Neke udičarke su takođe primetne po ekstremnom polnom dimorfizmu i polnoj simbiozi malog mužjaka sa mnogo većom ženkom, viđenom u podredu Ceratioidei. Kod ove vrste mužjaci mogu biti nekoliko redova veličine manji od ženki.
Udičarke su prisutne širom sveta. Neke su pelagijske (obitavaju udoaljeno od morskog dna), dok su druge bentoske (prebivaju u blizini morskog dna); neke žive u dubokom moru (npr. Ceratiidae ), dok druge nastanjuju kontinentalne pragove (npr. žabaste ribe Antennariidae i pripadnici porodice Lophiidae ). Pelagični oblici su uglavnom bočno komprimovani, dok su bentoški oblici često izuzetno dorsoventralno komprimovani (potisnuti), često sa velikim naviše usmerenim ustima.
Evolucija
Jedna filogenetička studija mitohondrijskog genoma je izvela zaključak da su se udičarke diverzifikovale u kratkom periodu od rane do središnje krede , pre između 130 i 100 miliona godina.
Klasifikacija
Ogcocephalus darwini
Prugasta udičarka (Antennarius striatus )
FishBase , Nelson, i Pič navode 18 familija, dok ITIS navodi samo 16. Sledeći taksoni su uređeni tako da su prikazani evolucioni odnosti.
Rodovi
Acentrophryne Regan, 1926
Allenichthys Pietsch, 1984
Antennarius Daudin, 1816
Antennatus Schultz, 1957
Bathychaunax Caruso, 1989
Bertella Pietsch, 1973
Borophryne Regan, 1925
Brachionichthys Bleeker, 1855
Bufoceratias Whitley, 1931
Caulophryne Goode and Bean, 1896
Centrophryne Regan and Trewavas, 1932
Ceratias Krøyer, 1845
Chaenophryne Regan, 1925
Chaunax Lowe, 1846
Chirophryne Regan and Trewavas, 1932
Coelophrys Brauer, 1902
Cryptopsaras Gill, 1883
Ctenochirichthys Regan and Trewavas, 1932
Danaphryne Bertelsen, 1951
Dermatias Smith and Radcliffe in Radcliffe, 1912
Dibranchus Peters, 1876
Diceratias Günther, 1887
Dolopichthys Garman, 1899
Echinophryne McCulloch and Waite, 1918
Gigantactis Brauer, 1902
Halicmetus Alcock, 1891
Halieutaea Valenciennes in Cuvier and Valenciennes, 1837
Halieutichthys Poey in Gill, 1863
Halieutopsis Garman, 1899
Haplophryne Regan, 1912
Himantolophus Reinhardt, 1837
Histiophryne Gill, 1863
Histrio Fischer, 1813
Kuiterichthys Pietsch, 1984
Lasiognathus Regan, 1925
Leptacanthichthys Regan and Trewavas, 1932
Linophryne Collett, 1886
Lophichthys Boeseman, 1964
Lophiocharon Whitley, 1933
Lophiodes Goode and Bean, 1896
Lophiomus Gill, 1883
Lophius Linnaeus, 1758
Lophodolos Lloyd, 1909
Malthopsis Alcock, 1891
Melanocetus Günther, 1864
Microlophichthys Burton, 1932
Neoceratias Pappenheim, 1914
Nudiantennarius Schultz, 1957
Ogcocephalus Fischer, 1813
Oneirodes Lütken, 1871
Pentherichthys Regan and Trewavas, 1932
Photocorynus Regan, 1925
Phyllophryne Pietsch, 1984
Phyllorhinichthys Pietsch, 1969
Puck Pietsch, 1978
Rhycherus Ogilby, 1907
Rhynchactis Regan, 1925
Robia Pietsch, 1979
Sladenia Regan, 1908
Solocisquama Bradbury, 1999
Spiniphryne Bertelsen, 1951
Sympterichthys Gill, 1878
Tathicarpus Ogilby, 1907
Tetrabrachium Günther, 1880
Thaumatichthys Smith and Radcliffe, 1912
Tyrannophryne Regan and Trewavas, 1932
Zalieutes Jordan and Evermann, 1896
Vrste
Acentrophryne longidens Regan, 1926
Allenichthys glauerti (Whitley, 1944)
Antennarius analis (Schultz, 1957)
Antennarius avalonis Jordan and Starks, 1907
Antennarius bermudensis Schultz, 1957
Antennarius biocellatus (Cuvier, 1817)
Antennarius coccineus (Lesson, 1831)
Antennarius commerson (Latreille, 1804)
Antennarius dorehensis Bleeker, 1859
Antennarius duescus Snyder, 1904
Antennarius hispidus (Bloch and Schneider, 1801)
Antennarius indicus Schultz, 1964
Antennarius maculatus (Desjardins, 1840)
Antennarius multiocellatus (Valenciennes in Cuvier and Valenciennes, 1837)
Antennarius nummifer (Cuvier, 1817)
Antennarius ocellatus (Bloch and Schneider, 1801)
Antennarius pardalis (Valenciennes in Cuvier and Valenciennes, 1837)
Antennarius pauciradiatus Schultz, 1957
Antennarius pictus (Shaw in Shaw and Nodder, 1794)
Antennarius radiosus Garman, 1896
Antennarius randalli Allen, 1970
Antennarius rosaceus Smith and Radcliffe in Radcliffe, 1912
Antennarius sanguineus Gill, 1863
Antennarius scriptissimus Jordan, 1902
Antennarius senegalensis Cadenat, 1959
Antennarius striatus (Shaw in Shaw and Nodder, 1794)
Antennatus flagellatus Ohnishi, Iwata and Hiramatsu, 1997
Antennatus linearis Randall and Holcom, 2001
Antennatus strigatus (Gill, 1863)
Antennatus tuberosus (Cuvier, 1817)
Bathychaunax coloratus (Garman, 1899)
Bathychaunax melanostomus Caruso, 1989
Bathychaunax roseus (Barbour, 1941)
Bertella idiomorpha Pietsch, 1973
Borophryne apogon Regan, 1925
Borophryne masculina Parr, 1934
Brachionichthys hirsutus (Lacepède, 1804)
Bufoceratias shaoi Pietsch, Ho and Chen, 2004
Bufoceratias thele (Uwate, 1979)
Bufoceratias wedli (Pietschmann, 1926)
Caulophryne jordani Goode and Bean, 1896
Caulophryne pelagica (Brauer, 1902)
Caulophryne pietschi Balushkin and Fedorov, 1985
Caulophryne polynema Regan, 1930
Centrophryne spinulosa Regan and Trewavas, 1932
Ceratias holboelli Krøyer, 1845
Ceratias tentaculatus (Norman, 1930)
Ceratias uranoscopus Murray in Thompson, 1877
Chaenophryne draco Beebe, 1932
Chaenophryne longiceps Regan, 1925
Chaenophryne melanorhabdus Regan and Trewavas, 1932
Chaenophryne ramifera Regan and Trewavas, 1932
Chaunax abei Le Danois, 1978
Chaunax breviradius Le Danois, 1978
Chaunax endeavouri Whitley, 1929
Chaunax fimbriatus Hilgendorf, 1879
Chaunax flammeus Le Danois, 1979
Chaunax latipunctatus Le Danois, 1984
Chaunax penicillatus McCulloch, 1915
Chaunax pictus Lowe, 1846
Chaunax stigmaeus Fowler, 1946
Chaunax suttkusi Caruso, 1989
Chaunax tosaensis Okamura and Oryuu in Okamura and Kitajima, 1984
Chaunax umbrinus Gilbert, 1905
Chirophryne xenolophus Regan and Trewavas, 1932
Coelophrys arca Smith and Radcliffe in Radcliffe, 1912
Coelophrys bradburyae Endo and Shinohara, 1999
Coelophrys brevicaudata Brauer, 1902
Coelophrys brevipes Smith and Radcliffe in Radcliffe, 1912
Coelophrys mollis Smith and Radcliffe in Radcliffe, 1912
Coelophrys oblonga Smith and Radcliffe in Radcliffe, 1912
Cryptopsaras couesii Gill, 1883
Ctenochirichthys longimanus Regan and Trewavas, 1932
Danaphryne nigrifilis (Regan and Trewavas, 1932)
Dermatias platynogaster Smith and Radcliffe in Radcliffe, 1912
Dibranchus accinctus Bradbury, 1999
Dibranchus atlanticus Peters, 1876
Dibranchus cracens Bradbury, McCosker and Long, 1999
Dibranchus discors Bradbury, McCosker and Long, 1999
Dibranchus erinaceus (Garman, 1899)
Dibranchus hystrix Garman, 1899
Dibranchus japonicus Amaoka and Toyoshima, 1981
Dibranchus nudivomer (Garman, 1899)
Dibranchus sparsus (Garman, 1899)
Dibranchus spinosus (Garman, 1899)
Dibranchus spongiosa (Gilbert, 1890)
Dibranchus tremendus Bradbury, 1999
Dibranchus velutinus Bradbury, 1999
Diceratias bispinosus (Günther, 1887)
Diceratias pileatus Uwate, 1979
Diceratias trilobus Balushkin and Fedorov, 1986
Dolopichthys allector Garman, 1899
Dolopichthys danae Regan, 1926
Dolopichthys dinema Pietsch, 1972
Dolopichthys jubatus Regan and Trewavas, 1932
Dolopichthys karsteni Leipertz and Pietsch, 1987
Dolopichthys longicornis Parr, 1927
Dolopichthys pullatus Regan and Trewavas, 1932
Echinophryne crassispina McCulloch and Waite, 1918
Echinophryne mitchellii (Morton, 1897)
Echinophryne reynoldsi Pietsch and Kuiter, 1984
Gigantactis balushkini Kharin, 1984
Gigantactis elsmani Bertelsen, Pietsch and Lavenberg, 1981
Gigantactis filibulbosus Fraser-Brunner, 1935
Gigantactis gargantua Bertelsen, Pietsch and Lavenberg, 1981
Gigantactis gibbsi Bertelsen, Pietsch and Lavenberg, 1981
Gigantactis golovani Bertelsen, Pietsch and Lavenberg, 1981
Gigantactis gracilicauda Regan, 1925
Gigantactis herwigi Bertelsen, Pietsch and Lavenberg, 1981
Gigantactis ios Bertelsen, Pietsch and Lavenberg, 1981
Gigantactis kreffti Bertelsen, Pietsch and Lavenberg, 1981
Gigantactis longicauda Bertelsen and Pietsch, 2002
Gigantactis longicirra Waterman, 1939
Gigantactis macronema Regan, 1925
Gigantactis meadi Bertelsen, Pietsch and Lavenberg, 1981
Gigantactis microdontis Bertelsen, Pietsch and Lavenberg, 1981
Gigantactis microphthalmus (Regan and Trewavas, 1932)
Gigantactis paxtoni Bertelsen, Pietsch and Lavenberg, 1981
Gigantactis perlatus Beebe and Crane, 1947
Gigantactis savagei Bertelsen, Pietsch and Lavenberg, 1981
Gigantactis vanhoeffeni Brauer, 1902
Gigantactis watermani Bertelsen, Pietsch and Lavenberg, 1981
Halicmetus reticulatus Smith and Radcliffe in Radcliffe, 1912
Halicmetus ruber Alcock, 1891
Halieutaea brevicauda Ogilby, 1910
Halieutaea coccinea Alcock, 1889
Halieutaea fitzsimonsi (Gilchrist and Thompson, 1916)
Halieutaea fumosa Alcock, 1894
Halieutaea hancocki Regan, 1908
Halieutaea indica Annandale and Jenkins, 1910
Halieutaea nigra Alcock, 1891
Halieutaea retifera Gilbert, 1905
Halieutaea stellata (Vahl, 1797)
Halieutichthys aculeatus (Mitchill, 1818)
Halieutopsis andriashevi Bradbury, 1988
Halieutopsis bathyoreos Bradbury, 1988
Halieutopsis galatea Bradbury, 1988
Halieutopsis ingerorum Bradbury, 1988
Halieutopsis micropa (Alcock, 1891)
Halieutopsis nasuta (Alcock, 1891)
Halieutopsis simula (Smith and Radcliffe in Radcliffe, 1912)
Halieutopsis stellifera (Smith and Radcliffe in Radcliffe, 1912)
Halieutopsis tumifrons Garman, 1899
Halieutopsis vermicularis Smith and Radcliffe in Radcliffe, 1912
Haplophryne mollis (Brauer, 1902)
Haplophryne triregium Whitley and Phillipps, 1939
Himantolophus albinares Maul, 1961
Himantolophus appelii (Clarke, 1878)
Himantolophus azurlucens Beebe and Crane, 1947
Himantolophus borealis Kharin, 1984
Himantolophus brevirostris (Regan, 1925)
Himantolophus compressus (Osório, 1912)
Himantolophus cornifer Bertelsen and Krefft, 1988
Himantolophus crinitus Bertelsen and Krefft, 1988
Himantolophus danae Regan and Trewavas, 1932
Himantolophus groenlandicus Reinhardt, 1837
Himantolophus macroceras Bertelsen and Krefft, 1988
Himantolophus macroceratoides Bertelsen and Krefft, 1988
Himantolophus mauli Bertelsen and Krefft, 1988
Himantolophus melanolophus Bertelsen and Krefft, 1988
Himantolophus multifurcatus Bertelsen and Krefft, 1988
Himantolophus nigricornis Bertelsen and Krefft, 1988
Himantolophus paucifilosus Bertelsen and Krefft, 1988
Himantolophus pseudalbinares Bertelsen and Krefft, 1988
Himantolophus sagamius (Tanaka, 1918)
Histiophryne bougainvilli (Valenciennes in Cuvier and Valenciennes, 1837)
Histiophryne cryptacanthus (Weber, 1913)
Histrio histrio (Linnaeus, 1758)
Kuiterichthys furcipilis (Cuvier, 1817)
Lasiognathus beebei Regan and Trewavas, 1932
Lasiognathus intermedius Bertelsen and Pietsch, 1996
Lasiognathus saccostoma Regan, 1925
Lasiognathus waltoni Nolan and Rosenblatt, 1975
Leptacanthichthys gracilispinis (Regan, 1925)
Linophryne algibarbata Waterman, 1939
Linophryne andersoni Gon, 1992
Linophryne arborifera Regan, 1925
Linophryne arcturi (Beebe, 1926)
Linophryne argyresca Regan and Trewavas, 1932
Linophryne bicornis Parr, 1927
Linophryne bipennata Bertelsen, 1982
Linophryne brevibarbata Beebe, 1932
Linophryne coronata Parr, 1927
Linophryne densiramus Imai, 1941
Linophryne digitopogon Balushkin and Trunov, 1988
Linophryne escaramosa Bertelsen, 1982
Linophryne indica (Brauer, 1902)
Linophryne lucifer Collett, 1886
Linophryne macrodon Regan, 1925
Linophryne maderensis Maul, 1961
Linophryne parini Bertelsen, 1980
Linophryne pennibarbata Bertelsen, 1980
Linophryne polypogon Regan, 1925
Linophryne quinqueramosa Beebe and Crane, 1947
Linophryne racemifera Regan and Trewavas, 1932
Linophryne sexfilis Bertelsen, 1973
Linophryne trewavasae Bertelsen, 1978
Lophichthys boschmai Boeseman, 1964
Lophiocharon hutchinsi Pietsch, 2004
Lophiocharon lithinostomus (Jordan and Richardson, 1908)
Lophiocharon trisignatus (Richardson, 1844)
Lophiodes abdituspinus Ni, Wu and Li, 1990
Lophiodes beroe Caruso, 1981
Lophiodes bruchius Caruso, 1981
Lophiodes caulinaris (Garman, 1899)
Lophiodes fimbriatus Saruwatari and Mochizuki, 1985
Lophiodes gracilimanus (Alcock, 1899)
Lophiodes infrabrunneus Smith and Radcliffe in Radcliffe, 1912
Lophiodes insidiator (Regan, 1921)
Lophiodes kempi (Norman, 1935)
Lophiodes miacanthus (Gilbert, 1905)
Lophiodes monodi (Le Danois, 1971)
Lophiodes mutilus (Alcock, 1894)
Lophiodes naresi (Günther, 1880)
Lophiodes reticulatus Caruso and Suttkus, 1979
Lophiodes spilurus (Garman, 1899)
Lophiomus setigerus (Vahl, 1797)
Lophius americanus Valenciennes in Cuvier and Valenciennes, 1837
Lophius budegassa Spinola, 1807
Lophius gastrophysus Miranda Ribeiro, 1915
Lophius litulon (Jordan, 1902)
Lophius lugubris Alcock, 1894
Lophius piscatorius Linnaeus, 1758
Lophius vomerinus Valenciennes in Cuvier and Valenciennes, 1837
Lophodolos acanthognathus Regan, 1925
Lophodolos indicus Lloyd, 1909
Malthopsis annulifera Tanaka, 1908
Malthopsis gnoma Bradbury, 1998
Malthopsis jordani Gilbert, 1905
Malthopsis lutea Alcock, 1891
Malthopsis mitrigera Gilbert and Cramer, 1897
Malthopsis tiarella Jordan, 1902
Melanocetus eustalus Pietsch and Van Duzer, 1980
Melanocetus johnsonii Günther, 1864
Melanocetus murrayi Günther, 1887
Melanocetus niger Regan, 1925
Melanocetus rossi Balushkin and Fedorov, 1981
Microlophichthys andracanthus Bertelsen, 1951
Microlophichthys microlophus (Regan, 1925)
Neoceratias spinifer Pappenheim, 1914
Nudiantennarius subteres (Smith and Radcliffe in Radcliffe, 1912)
Ogcocephalus corniger Bradbury, 1980
Ogcocephalus cubifrons (Richardson, 1836)
Ogcocephalus darwini Hubbs, 1958
Ogcocephalus declivirostris Bradbury, 1980
Ogcocephalus nasutus (Cuvier, 1829)
Ogcocephalus notatus (Valenciennes in Cuvier and Valenciennes, 1837)
Ogcocephalus pantostictus Bradbury, 1980
Ogcocephalus parvus Longley and Hildebrand, 1940
Ogcocephalus porrectus Garman, 1899
Ogcocephalus pumilus Bradbury, 1980
Ogcocephalus rostellum Bradbury, 1980
Ogcocephalus vespertilio (Linnaeus, 1758)
Oneirodes acanthias (Gilbert, 1915)
Oneirodes alius Seigel and Pietsch, 1978
Oneirodes anisacanthus (Regan, 1925)
Oneirodes appendixus Ni and Xu in Ni, 1988
Oneirodes basili Pietsch, 1974
Oneirodes bradburyae Grey, 1956
Oneirodes bulbosus Chapman, 1939
Oneirodes carlsbergi (Regan and Trewavas, 1932)
Oneirodes clarkei Swinney and Pietsch, 1988
Oneirodes cristatus (Regan and Trewavas, 1932)
Oneirodes dicromischus Pietsch, 1974
Oneirodes epithales Orr, 1991
Oneirodes eschrichtii Lütken, 1871
Oneirodes flagellifer (Regan and Trewavas, 1932)
Oneirodes haplonema Stewart and Pietsch, 1998
Oneirodes heteronema (Regan and Trewavas, 1932)
Oneirodes kreffti Pietsch, 1974
Oneirodes luetkeni (Regan, 1925)
Oneirodes macronema (Regan and Trewavas, 1932)
Oneirodes macrosteus Pietsch, 1974
Oneirodes melanocauda Bertelsen, 1951
Oneirodes micronema Grobecker, 1978
Oneirodes mirus (Regan and Trewavas, 1932)
Oneirodes myrionemus Pietsch, 1974
Oneirodes notius Pietsch, 1974
Oneirodes pietschi Ho and Shao, 2004
Oneirodes plagionema Pietsch and Seigel, 1980
Oneirodes posti Bertelsen and Grobecker, 1980
Oneirodes pterurus Pietsch and Seigel, 1980
Oneirodes rosenblatti Pietsch, 1974
Oneirodes sabex Pietsch and Seigel, 1980
Oneirodes schistonema Pietsch and Seigel, 1980
Oneirodes schmidti (Regan and Trewavas, 1932)
Oneirodes theodoritissieri Belloc, 1938
Oneirodes thompsoni (Schultz, 1934)
Oneirodes thysanema Pietsch and Seigel, 1980
Oneirodes whitleyi Bertelsen and Pietsch, 1983
Pentherichthys atratus (Regan and Trewavas, 1932)
Pentherichthys venustus (Regan and Trewavas, 1932)
Photocorynus spiniceps Regan, 1925
Phyllophryne scortea (McCulloch and Waite, 1918)
Phyllorhinichthys micractis Pietsch, 1969
Puck pinnata Pietsch, 1978
Rhycherus filamentosus (Castelnau, 1872)
Rhycherus gloveri Pietsch, 1984
Rhynchactis leptonema Regan, 1925
Rhynchactis macrothrix Bertelsen and Pietsch, 1998
Rhynchactis microthrix Bertelsen and Pietsch, 1998
Robia legula Pietsch, 1979
Sladenia remiger Smith and Radcliffe in Radcliffe, 1912
Sladenia shaefersi Caruso and Bullis, 1976
Solocisquama carinata Bradbury, 1999
Solocisquama erythrina (Gilbert, 1905)
Solocisquama stellulata (Gilbert, 1905)
Spiniphryne gladisfenae (Beebe, 1932)
Sympterichthys politus (Richardson, 1844)
Sympterichthys unipennis (Cuvier, 1817)
Sympterichthys verrucosus McCulloch and Waite, 1918
Tathicarpus butleri Ogilby, 1907
Tetrabrachium ocellatum Günther, 1880
Thaumatichthys axeli (Bruun, 1953)
Thaumatichthys binghami Parr, 1927
Thaumatichthys pagidostomus Smith and Radcliffe, 1912
Tyrannophryne pugnax Regan and Trewavas, 1932
Zalieutes elater (Jordan and Gilbert, 1882)
Zalieutes mcgintyi (Fowler, 1952)
Hronologija
Udičarke se javljaju u fosilnom zapisu u sledećem redosledu:
Reference
Literatura
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