Species in mammal family Canidae
10 of the 13 extant canid genera left-to-right, top-to-bottom: Canis , Cuon , Lycaon , Cerdocyon , Chrysocyon , Speothos , Vulpes , Nyctereutes , Otocyon , and Urocyon
Canidae is a family of mammals in the order Carnivora , which includes domestic dogs , wolves , coyotes , foxes , jackals , dingoes , and many other extant and extinct dog-like mammals. A member of this family is called a canid; all extant species are a part of a single subfamily, Caninae , and are called canines. They are found on all continents except Antarctica, having arrived independently or accompanied human beings over extended periods of time. Canids vary in size, including tails, from the 2 meter (6 ft 7 in) wolf to the 46 cm (18 in) fennec fox . Population sizes range from the Falkland Islands wolf , extinct since 1876, to the domestic dog, which has a worldwide population of over 1 billion. The body forms of canids are similar, typically having long muzzles, upright ears, teeth adapted for cracking bones and slicing flesh, long legs, and bushy tails. Most species are social animals, living together in family units or small groups and behaving cooperatively. Typically, only the dominant pair in a group breeds, and a litter of young is reared annually in an underground den. Canids communicate by scent signals and vocalizations. One canid, the domestic dog, entered into a partnership with humans at least 14,000 years ago and today remains one of the most widely kept domestic animals.
The 13 extant genera and 37 species of Caninae are primarily split into two tribes : Canini, which includes 11 genera and 19 species, comprising the wolf-like Canina subtribe and the South American Cerdocyonina subtribe; and Vulpini, the fox-like canids, comprising 3 genera and 15 species. Not included in either tribe is the genus Urocyon , which includes 2 species, mainly comprising the gray fox and believed to be basal to the family. Additionally, one genus in Canini, Dusicyon , was composed of two recently extinct species, with Dusicyon avus going extinct around 400 years ago and the Falkland Islands wolf going extinct in 1876.
In addition to the extant Caninae, Canidae contains two extinct subfamilies designated as Hesperocyoninae and Borophaginae . Extinct species have also been placed into Caninae, in both extant and extinct genera; at least 80 extinct Caninae species have been found, as well as over 70 species in Borophaginae and nearly 30 in Hesperocyoninae, though due to ongoing research and discoveries the exact number and categorization is not fixed. The earliest canids found belong to Hesperocyoninae, and are believed to have diverged from the existing Caniformia suborder around 37 million years ago.
Conventions
Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species . Range maps are provided wherever possible; if a range map is not available, a description of the canid's range is provided. Ranges are based on the IUCN Red List for that species, unless otherwise noted. All extinct species (or subspecies) listed alongside extant species went extinct after 1500 CE, and are indicated by a dagger symbol:
"†".
Classification
The family Canidae consists of 37 extant species belonging to 13 genera and divided into 194 extant subspecies, as well the extinct genus Dusicyon , comprising two extinct species, and 13 extinct wolf subspecies, which are the only canid species to go extinct since prehistoric times. This does not include hybrid species (such as wolfdogs or coywolves ) or extinct prehistoric species (such as the dire wolf or Epicyon ). Modern molecular studies indicate that the 13 genera can be grouped into 3 tribes or clades .
Canids
The following classification is based on the taxonomy described by Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis , such as the promotion of the African golden wolf to a separate species from the golden jackal , and splitting out the Lupulella genus from Canis . Range maps are based on IUCN range data. There are several additional proposals which are disputed, such as the promotion of the red wolf and eastern wolf as species from subspecies of the wolf , which are marked with a "(debated)" tag.
Subfamily Caninae
Tribe Canini
Genus Atelocynus – Cabrera , 1940 – one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Short-eared dog
A. microtis (Cabrera , 1940)
A. m. microtis
A. m. sclateri
Western Amazon rainforest in South America
Size : 72–100 cm (28–39 in) long, plus 24–35 cm (9–14 in) tailHabitat : Wetlands, forest, and savannaDiet : Fish, insects, and small mammals, as well as fruit, birds, and crabs
NT
Unknown
Genus Canis – Linnaeus , 1758 – six species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
African wolf
C. lupaster Hemprich and Ehrenberg , 1832
North and northeastern Africa
Size : 100 cm (39 in) long, plus 20 cm (8 in) tailHabitat : Grassland, shrubland, and savannaDiet : Wild boar and livestock, as well as other mammals and fruit
LC
Unknown
Coyote
C. latrans Say , 1823
C. l. cagottis (Mexican coyote)
C. l. clepticus (San Pedro Martir coyote)
C. l. dickeyi (El Salvador coyote)
C. l. frustor (Southeastern coyote)
C. l. goldmani (Belize coyote)
C. l. hondurensis (Honduras coyote)
C. l. impavidus (Durango coyote)
C. l. incolatus (Northern coyote)
C. l. jamesi (Tiburón Island coyote)
C. l. latrans (Plains coyote )
C. l. lestes (Mountain coyote)
C. l. mearnsi (Mearns' coyote )
C. l. microdon (Lower Rio Grande coyote)
C. l. ochropus (California valley coyote)
C. l. peninsulae (Peninsula coyote)
C. l. texensis (Texas plains coyote)
C. l. thamnos (Northeastern coyote )
C. l. umpquensis (Northwest coast coyote)
C. l. vigilis (Colima coyote)
North America
Size : 100–135 cm (39–53 in) long, plus 40 cm (16 in) tailHabitat : Forest, desert, shrubland, and grasslandDiet : Wide variety of foods, including both small and large mammals, fruit, and insects
LC
1 million+
Dog
C. familiaris Linnaeus , 1758
Worldwide
Size : Varies by breedHabitat : DomesticatedDiet : Varied
NE
1 billion
Ethiopian wolf
C. simensis Rüppell , 1840
C. s. citernii (Southern Ethiopian wolf)
C. s. simensis (Northern Ethiopian wolf)
Ethiopian Highlands
Size : 84–100 cm (33–39 in) long, plus 27–40 cm (11–16 in) tailHabitat : Inland wetlands, grassland, shrubland, and rocky areasDiet : Rodents as well as small mammals
EN
200
Golden jackal
C. aureus Linnaeus , 1758
C. a. aureus (Persian jackal)
C. a. cruesemanni (Indochinese jackal)
C. a. indicus (Indian jackal)
C. a. moreoticus (European jackal )
C. a. naria (Sri Lankan jackal )
C. a. syriacus (Syrian jackal)
Eastern Europe, Middle East, and southern Asia
Size : 60–132 cm (24–52 in) long, plus 20–30 cm (8–12 in) tailHabitat : Forest, grassland, shrubland, and savannaDiet : Wide variety of foods, including small to large mammals, birds, fish, fruit, and insects
LC
Unknown, but at least 150,000
Wolf
C. lupus Linnaeus , 1758
Eurasia and northern North America
Size : 105–160 cm (41–63 in) long, plus 29–50 cm (11–20 in) tailHabitat : Forest, desert, rocky areas, shrubland, grassland, and inland wetlandsDiet : Large ungulates , as well as small animals, carrion, and berries
LC
300,000
Genus Cerdocyon – C. E. H. Smith , 1839 – one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Crab-eating fox
C. thous (Linnaeus , 1766)
C. t. aquilus
C. t. azarae
C. t. entrerianus
C. t. germanus
C. t. thous
Eastern and northern South America
Size : 64 cm (25 in) long, plus 28 cm (11 in) tailHabitat : Forest, savanna, shrubland, grassland, and inland wetlandsDiet : Crabs and insects, as well as rodents, birds, turtles, eggs, fruit, and carrion
LC
Unknown
Genus Chrysocyon – C. E. H. Smith , 1839 – one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Maned wolf
C. brachyurus (Illiger , 1815)
Central South America
Size : 100–130 cm (39–51 in) long, plus 45 cm (18 in) tailHabitat : Forest, wetlands, grassland, shrubland, and savannaDiet : Fruit, arthropods, and small and medium vertebrates
NT
17,000
Genus Cuon – Hodgson , 1838 – one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Dhole
C. alpinus (Pallas , 1811)
Southeast Asia
Size : 90 cm (35 in) long, plus 40–45 cm (16–18 in) tailHabitat : Forest, grassland, and shrublandDiet : Ungulates, as well as small rodents and hares
EN
1,000–2,200
Genus Dusicyon † – C. E. H. Smith , 1839 – two species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Falkland Islands wolf †
D. australis (Kerr , 1792)
Falkland Islands at tip of South America
Size : UnknownHabitat : Grassland and shrublandDiet : Unknown
EX
0
South American fox †
D. avus (Burmeister , 1866)
Southern South America
Size : UnknownHabitat : Grassland and shrublandDiet : Unknown
EX
0
Genus Lupulella – Hilzheimer , 1906 – two species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Black-backed jackal
L. mesomelas (Schreber , 1775)
L. m. mesomelas (Cape black-backed jackal)
L. m. schmidti (East African black-backed jackal)
Southern Africa and eastern Africa
Size : 60–95 cm (24–37 in) long, plus 16–40 cm (6–16 in) tailHabitat : Marine intertidal, forest, desert, grassland, shrubland, and savannaDiet : Small to medium-sized mammals and birds
LC
Unknown
Side-striped jackal
L. adustus (Sundevall , 1847)
L. a. adustus (Sundevall's side-striped jackal)
L. a. bweha
L. a. centralis
L. a. grayi
L. a. kaffensis (Kaffa side-striped jackal)
L. a. lateralis
L. a. notatus (East African side-striped jackal)
Central Africa
Size : 69–81 cm (27–32 in) long, plus 30–41 cm (12–16 in) tailHabitat : Forest, shrubland, savanna, grassland, and inland wetlandsDiet : Small to medium-sized mammals and fruit, as well as birds, insects, grass, and carrion
LC
3 million
Genus Lycalopex – Burmeister , 1854 – six species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Culpeo
L. culpeo (Molina , 1782)
L. c. andinus
L. c. culpaeus
L. c. lycoides
L. c. magellanicus
L. c. reissii
L. c. smithersi
Western South America
Size : 95–132 cm (37–52 in) long, plus 32–44 cm (13–17 in) tailHabitat : Forest, rocky areas, grassland, shrubland, and savannaDiet : Rodents and lagomorphs , as well as livestock and guanacos
LC
Unknown
Darwin's fox
L. fulvipes (Martin , 1837)
Limited areas in southern Chile
Size : 48–59 cm (19–23 in) long, plus 18–26 cm (7–10 in) tailHabitat : Forest and shrublandDiet : Small mammals, insects, crabs, and fruit
EN
600-2,500
Hoary fox
L. vetulus (Lund , 1842)
South-central Brazil
Size : 49–71 cm (19–28 in) long, plus 25–38 cm (10–15 in) tailHabitat : SavannaDiet : Insects, as well as small rodents, birds, reptiles, and fruit
LC
Unknown
Pampas fox
L. gymnocercus (Waldheim , 1814)
L. g. antiquus
L. g. domeykoanus
L. g. gracilis
L. g. gymnocercus
L. g. maulinicus
Southern South America
Size : 51–74 cm (20–29 in) long, plus 25–41 cm (10–16 in) tailHabitat : Forest, shrubland, and savannaDiet : Small rodents, hares, birds, insects, and fruit, as well as carrion
LC
Unknown
Sechuran fox
L. sechurae (Thomas , 1900)
Sechura Desert in southwestern Ecuador and northwestern Peru
Size : 50–78 cm (20–31 in) long, plus 27–34 cm (11–13 in) tailHabitat : Forest, desert, grassland, and shrublandDiet : Fruit and seeds, as well as small rodents, birds, reptiles, insects, scorpions, and carrion
NT
15,000
South American gray fox
L. griseus (Gray , 1837)
Southern South America
Size : 50–66 cm (20–26 in) long, plus 12–34 cm (5–13 in) tailHabitat : Forest, grassland, and shrublandDiet : Small rodents, hares, and carrion
LC
Unknown
Genus Lycaon – Brookes , 1827 – one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
African wild dog
L. pictus (Temminck , 1820)
Scattered areas of Africa. Extant regions in red; probably extant region in yellow.
Size : 76–112 cm (30–44 in) long, plus 30–42 cm (12–17 in) tailHabitat : Forest, grassland, shrubland, savanna, and desertDiet : Medium-sized antelope
EN
1,400
Genus Speothos – Lund , 1839 – one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Bush dog
S. venaticus (Lund , 1842)
S. v. panamensis (Panamanian bush dog)
S. v. venaticus (South American bush dog)
S. v. wingei (Southern bush dog)
Northern South America
Size : 57–75 cm (22–30 in) long, plus 12–15 cm (5–6 in) tailHabitat : Shrubland, forest, grassland, and savannaDiet : Small and medium mammals, as well as birds, reptiles, and fruit
NT
15,000
Tribe Vulpini
Genus Nyctereutes – Temminck , 1839 – two species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Common raccoon dog
N. procyonoides (Gray , 1834)
N. p. procyonoides (Chinese raccoon dog)
N. p. koreensis (Korean raccoon dog)
N. p. orestes (Yunnan raccoon dog)
N. p. ussuriensis (Ussuri raccoon dog)
Mainland Eastern Asia, introduced to Central and Eastern Europe (note: map includes range of N. viverrinus )
Size : 49–71 cm (19–28 in) long, plus 15–23 cm (6–9 in) tailHabitat : Forest, grassland, and shrublandDiet : Insects, rodents, amphibians, birds, fish, and reptiles, as well as fruit, nuts, and berries
LC
Unknown, but at least 1.5 million in fur farms
Japanese raccoon dog
N. viverrinus (Temminck, 1838)
Japan
Size : 49–71 cm (19–28 in) long, plus 15–23 cm (6–9 in) tailHabitat : Forest, grassland, and shrublandDiet : Insects, rodents, amphibians, birds, fish, and reptiles, as well as fruit, nuts, and berries
NE
Unknown
Genus Otocyon – Müller , 1835 – one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Bat-eared fox
O. megalotis (Desmarest , 1822)
O. m. megalotis
O. m. virgatus
Southern and Eastern Africa
Size : 46–61 cm (18–24 in) long, plus 23–34 cm (9–13 in) tailHabitat : Grassland, shrubland, and savannaDiet : Harvester termites as well as other arthropods
LC
Unknown
Genus Vulpes – Frisch , 1775 – twelve species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Arctic fox
V. lagopus (Linnaeus , 1758)
V. l. lagopus (Common Arctic Fox)
V. l. beringensis (Bering Islands Arctic fox)
V. l. foragoapusis (Greenland Arctic fox)
V. l. fuliginosus (Iceland Arctic fox)
V. l. pribilofensis (Pribilof Islands Arctic fox)
Arctic North America and Eurasia
Size : 50–75 cm (20–30 in) long, plus 25–43 cm (10–17 in) tailHabitat : GrasslandDiet : Lemmings, as well as other rodents, birds, and reindeer
LC
Unknown
Bengal fox
V. bengalensis (Shaw , 1800)
India
Size : 39–58 cm (15–23 in) long, plus 25–32 cm (10–13 in) tailHabitat : Grassland and shrublandDiet : Arthropods, rodents, reptiles, fruit, and birds
LC
Unknown
Blanford's fox
V. cana Blanford , 1877
The Middle East and Central Asia
Size : 34–47 cm (13–19 in) long, plus 26–36 cm (10–14 in) tailHabitat : Desert and rocky areasDiet : Fruit and insects
LC
Unknown
Cape fox
V. chama (A Smith , 1833)
Southern Africa
Size : 45–61 cm (18–24 in) long, plus 25–41 cm (10–16 in) tailHabitat : Rocky areas, grassland, shrubland, and savannaDiet : Fruit and insects
LC
20,000
Corsac fox
V. corsac (Linnaeus , 1768)
V. c. corsac
V. c. kalmykorum
V. c. turkmenicus
Central Asia
Size : 45–60 cm (18–24 in) long, plus 19–34 cm (7–13 in) tailHabitat : Desert, grassland, and shrublandDiet : Insects and small rodents
LC
Unknown
Fennec fox
V. zerda (Zimmermann , 1780)
Northern Africa
Size : 33–40 cm (13–16 in) long, plus 13–23 cm (5–9 in) tailHabitat : Desert and marine coastal/supratidalDiet : Rodents, insects, birds, eggs, and rabbits
LC
Unknown
Kit fox
V. macrotis Merriam , 1888
Western North America
Size : 46–54 cm (18–21 in) long, plus 25–34 cm (10–13 in) tailHabitat : Shrubland, savanna, and grasslandDiet : Rodents, rabbits, invertebrates, birds, lizards, and snakes
LC
Unknown
Pale fox
V. pallida (Cretzschmar , 1827)
V. p. cyrenaica
V. p. edwardsi
V. p. harterti
V. p. oertzeni
V. p. pallida
Upper middle Africa
Size : 38–55 cm (15–22 in) long, plus 23–29 cm (9–11 in) tailHabitat : Desert, grassland, shrubland, and savannaDiet : Plants and berries as well as rodents, reptiles, and insects
LC
10,000–100,000
Rüppell's fox
V. rueppellii (Schinz , 1825)
Northern Africa and the Middle East
Size : 35–56 cm (14–22 in) long, plus 25–39 cm (10–15 in) tailHabitat : Desert, shrubland, and marine coastal/supratidalDiet : Small mammals, lizards, birds, and insects, as well as fruit and succulents
LC
Unknown
Red fox
V. vulpes (Linnaeus , 1758)
V. v. abietorum (British Columbian fox)
V. v. alascensis (Northern Alaskan fox)
V. v. alpherakyi (Eastern Trans-Caucasian fox)
V. v. anatolica (Anatolian fox)
V. v. arabica (Arabian red fox )
V. v. atlantica (Atlas fox)
V. v. bangsi (Labrador fox)
V. v. barbara (Barbary fox)
V. v. beringiana (Anadyr fox)
V. v. cascadensis (Cascade red fox )
V. v. caucasica (North Caucasian fox)
V. v. crucigera (European fox)
V. v. daurica (Trans-Baikal fox)
V. v. deletrix (Newfoundland fox)
V. v. dolichocrania (Ussuri fox)
V. v. dorsalis
V. v. lavescens (Turkmenian fox )
V. v. fulvus (American red fox )
V. v. harrimani (Afghan red fox)
V. v. hoole (Southern Chinese fox)
V. v. ichnusae (Sardinian fox)
V. v. indutus (Cyprus fox)
V. v. jakutensis (Yakutsk fox)
V. v. japonica (Japanese fox)
V. v. karagan (Karaganka fox)
V. v. kenaiensis (Kenai Peninsula fox)
V. v. kurdistanica (Trans-Caucasian fox )
V. v. macroura (Wasatch Mountains fox)
V. v. montana (Hill fox)
V. v. necator (Sierra Nevada red fox )
V. v. niloticus (Nile fox)
V. v. ochroxantha (Turkestan fox)
V. v. palaestina (Palestinian fox)
V. v. peculiosa (Korean fox )
V. v. pusilla (White-footed fox )
V. v. regalis (Northern plains fox)
V. v. rubricosa (Nova Scotia fox)
V. v. schrencki (Sakhalin fox)
V. v. silacea (Iberian fox)
V. v. splendidissima (Kuril Islands fox)
V. v. stepensis (Steppe red fox)
V. v. tobolica (Tobol'sk fox)
V. v. tschiliensis (Northern Chinese fox)
V. v. vulpes (Scandinavian red fox)
North America, Europe, Asia, and Australia
Size : 62–72 cm (24–28 in) long, plus 40 cm (16 in) tailHabitat : Shrubland, grassland, inland wetlands, forest, and desertDiet : Small rodents, as well as birds, larger mammals, reptiles, insects, and fish
LC
Unknown
Swift fox
V. velox (Say , 1823)
Western grasslands of North America
Size : 48–54 cm (19–21 in) long, plus 25–34 cm (10–13 in) tailHabitat : GrasslandDiet : Rabbits, mice, ground squirrels, birds, insects and lizards, as well as grasses and fruit
LC
Unknown
Tibetan fox
V. ferrilata Hodgson , 1842
High plateaus in Nepal and western China
Size : 49–70 cm (19–28 in) long, plus 22–29 cm (9–11 in) tailHabitat : Desert, rocky areas, grassland, and shrublandDiet : Pikas , as well as carrion and other small mammals
LC
Unknown
Urocyon
Genus Urocyon – Baird , 1857 – two species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Gray fox
U. cinereoargenteus (Schreber , 1775)
U. c. borealis
U. c. californicus
U. c. cinereoargenteus
U. c. costaricensis
U. c. floridanus
U. c. fraterculus
U. c. furvus
U. c. guatemalae
U. c. madrensis
U. c. nigrirostris
U. c. ocythous
U. c. orinomus
U. c. peninsularis
U. c. scottii
U. c. townsendi
U. c. venezuelae
North America and Central America
Size : 53–66 cm (21–26 in) long, plus 28–44 cm (11–17 in) tailHabitat : Forest and shrublandDiet : Rabbits, voles, shrews, and birds, as well as insects and fruit
LC
Unknown
Island fox
U. littoralis (Baird , 1857)
U. l. catalinae
U. l. clementae
U. l. dickeyi
U. l. littoralis
U. l. santacruzae
U. l. santarosae
Channel Islands of California
Size : 46–63 cm (18–25 in) long, plus 12–32 cm (5–13 in) tailHabitat : Marine intertidal, forest, grassland, and shrublandDiet : Fruit, insects, birds, eggs, crabs, lizards, and small mammals
NT
4,000
Notes
^ a b c d e f g h i j k l m n Population figures rounded to the nearest hundred. Population trends as described by the International Union for Conservation of Nature .
^ The Falklands Island wolf is believed to have been driven extinct in 1876
^ The South American fox is believed to have gone extinct sometime between 1454 and 1626
References
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^ Leite Pitman, Renata; S. R. Williams, Robert (2004). Sillero-Zubiri, C.; Gingsberg, J. R.; Macdonald, D. W. (eds.). Canids: Species status and conservation (2004 ed.). International Union for Conservation of Nature . pp. 26–31.
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